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We are proud of our alumni! Here is an chronological roster of our graduates and current professional details.
All dissertations are available from
UMI Dissertation Services, or by interlibrary loan.

  Dr. Christopher Beard
  • Foundation Distinguished Professor of Ecology & Evolutionary Biology, and Senior Curator in the Biodiversity Institute, The University of Kansas, Lawrence, KS
  • Ph.D., 1989 - Postcranial anatomy, locomotor adaptations, and paleoecology of early Cenozoic Plesiadapidae, Paromomyidae, and Micromomyidae (Eutheria, Dermoptera)
  • Associate Editor, Journal of Human Evolution (Academic Press)
  • MacArthur Fellow, 2000
  • E-mail: chris.beard(at)
  • Homepage:

Dr. Audrone Biknevicius
  • Associate Professor and Chair, Department of Biomedical Sciences, Department of Biomedical Sciences,College of Osteopathic Medicine, Ohio University, Athens, OH
  • Ph.D., 1990 - Biomechanical scaling of the mandibular corpus in carnivores
  • E-mail: biknevic(at)

Dr. Carol Ward
  • Professor, Department of Anthropology and Department of Pathology and Anatomical Sciences, University of Missouri, Columbia, MO
  • Ph.D., 1991 - Functional anatomy of the lower back and pelvis of the Miocene hominoid Proconsul nyanzae from Mfangano Island, Kenya
  • E-mail: WardCV(at)
  • Homepage:

Dr. Hannah Grausz
  • Associate Director, MetroHealth/Cleveland Clinic Foundation Emergency Medicine Residency and Associate Staff Physician, Cleveland Clinic Foundation, Cleveland, OH
  • Ph.D., 1992 - Growth of the human perinatal craniofacial skeleton characterized in three dimensions
  • M.D., 1996
  • E-mail: grauszh(at)

Dr. Lawrence M. Witmer
  • Professor of Anatomy, Chang Ying-Chien Professor of Paleontology, Department of Biomedical Sciences, College of Osteopathic Medicine, Ohio University, Athens, OH
  • Ph.D., 1992 - Ontogeny, phylogeny, and air sacs: The importance of soft-tissue inferences in the interpretation of facial evolution in Archosauria
  • E-mail: witmer1(at)
  • Homepage:

Dr. Jacqueline Runestad-Connour
  • Associate Professor, Department of Biology, University of Findlay, Findlay, OH
  • Ph.D., 1994 - Humeral and femoral diaphyseal cross-sectional geometry and articular dimensions in Prosimii and Platyrrhini (Primates) with application for reconstruction of body mass and locomotor behavior in Adapidae (Primates: Eocene)
  • E-mail: jackie_runestad(at)

Dr. Ronald E. Heinrich
  • Most recent academic address: Instructor, Division of Biological Sciences, University of Kansas, Lawrence, KS
  • Ph.D., 1996 - Functional morphology and body size of Early Tertiary Miacoidea (Mammalia, Carnivora)
  • E-mail: reheinrich(at)

Dr. Maureen O'Leary

Dr. Katherine Rafferty
  • Senior Lecturer, Department of Orthodontics, University of Washington School of Dentistry
  • Ph.D., 1996 - Joint design in primates: external and subarticular properties in relation to body size and locomotor behavior
  • E-mail: kraff(at)

Dr. Katherine Coffing
  • Most recent academic address: Lecturer, Department of Anthropology, University of California, Los Angeles, CA
  • Ph.D., 1998 - The metacarpals of Australopithecus afarensis: locomotor and behavioral implications of cross-sectional geometry
  • E-mail: kcoffing(at)

Dr. Naoko Egi

Dr. Mason Meers
  • Professor and Chair of Biology, Department of Biology, The University of Tampa, FL
  • Ph.D., 2000 - Evolution of the crocodylian forelimb: anatomy, biomechanics and functional morphology
  • E-mail: mmeers(at)

Dr. Gail Krovitz
  • Director of Academic Training & Consulting, Pearson eCollege, Denver, CO
  • Ph.D., 2001 - Three-dimensional comparisons of craniofacial morphologyand growth patterns in Neandertals and modern humans
  • E-mail: gailk(at)

Dr. Mary Silcox
  • Associate Professor of Anthropology at University of Toronto Scarbrough, Canada
  • Ph.D., 2001 - A phylogenetic analysis of the Plesiadapiformes and their relationship to Euprimates and other Archontans
  • E-mail: msilcox(at)

Dr. Brenda Chinnery
  • Most recent academic address: Lecturer, School of Biological Sciences, University of Texas at Austin, Austin, TX
  • Ph.D., 2002 - Morphometric analysis of evolution and growth in the Ceratopsian postcranial skeleton
  • E-mail: brendachinnery(at)

Dr. Yizheng Li
  • Staff Scientist, Wyeth Research, Cambridge, MA
  • Ph.D., 2002 - Postnatal development of pelvic sexual dimorphism in four anthropoid primates
  • E-mail: yli(at)

Dr. Kristina Aldridge
  • Assistant Professor, Department of Pathology & Anatomical Sciences, University of Missouri-Columbia, Columbia, MO
  • Ph.D., 2003 - Organization of the human brain: development, variability, and evolution
  • E-mail: aldridgek(at)

Dr. Anita Hettena
  • Biology Co-Chair and Assistant Professor of Biology, San Diego City College, San Diego, CA
  • Ph.D., 2003 - A three-dimensional analysis of age-related change in the adult craniofacial skeleton
  • E-mail: anitahettena(at)

Dr. Valerie Burke DeLeon
  • Assistant Professor at the Department of Anthropology, University of Florida. Adjunct Professor, Center for Functional Anatomy & Evolution, Johns Hopkins University School of Medicine, Baltimore, MD
  • Ph.D., 2004 - Fluctuating asymmetry in the human craniofacial skeleton: effects of sexual dimorphism, stress, and developmental anomalies
  • E-mail: vdeleon(at)

Dr. François Therrien
  • Curator of Dinosaur Palaeoecology, Royal Tyrrell Museum and Adjunct Assistant Professor, Department of Geoscience, University of Calgary, Canada
  • Ph.D., 2004 - Paleoenvironmental reconstruction of the Late Cretaceous (Maastrichtian) dinosaur-bearing formations of Romania
  • E-mail: Francois.Therrien(at)
  • Homepage:

Dr. Jay Mussell
  • Assistant Professor in the Department of Cell Biology and Anatomy, Louisiana State University Health Sciences Center New Orleans, LA.
  • Ph.D., 2004 - A reexamination of Lipotyphla and Afrotheria using both molecular and morphological analyses
  • E-mail: jmusse(at)

Dr. Ann C. Zumwalt
  • Assistant Professor in the Department of Anatomy & Neurobiology, Boston University School of Medicine, Boston, MA
  • Ph.D., 2004 - The effect of endurance exercise on the morphology of muscle attachment sites: An experimental study in sheep (Ovis aries)
  • E-mail: azumwalt(at)

Dr. Amy E. Chew
  • Associate Professor of Anatomy, College of Osteopathic Medicine of the Pacific, Western University of Health Sciences, Pomona, CA
  • Ph.D., 2005 - Biostratigraphy, Paleoecology and synchronized evolution in the Early Eocene mammalian fauna of the Central Bighorn Basin, Wyoming
  • E-mail: achew(at)

Dr. Benjamin M. Auerbach
  • Associate Professor, Department of Anthropology, The University of Tennessee, Knoxville, TN
  • Ph.D., 2007 - Skeletal variation in the New World during the Holocene: effects of climate and subsistence across geography and time
  • E-mail: auerbach(at)

Dr. Jason M. Organ
  • Assistant Professor, Department of Anatomy and Cell Biology, Indiana University School of Medicine, Indianapolis, IN
  • Ph.D., 2007 - The functional anatomy of prehensile and nonprehensile tails of the Platyrrhini (Primates) and Procyonidae (Carnivora)
  • E-mail: jorgan(at)
Dr. Matthew O'Neill
  • Research Instructor, Department of Anatomical Sciences, Stony Brook University
  • Ph.D., 2008 - The Structural Basis of Locomotor Cost: Gait, Mechanics and Limb Design in Ringtailed Lemurs (Lemur catta)
  • E-mail: matthew.oneill(at)

Dr. Shawn P. Zack
  • Assistant Professor of Anatomy, University of Arizona College of Medicine-Phoenix
  • Ph.D., 2009 - The phylogeny of eutherian mammals: a new analysis emphasizing dental and postcranial morphology of Paleogene taxa
  • E-mail: spzack(at)

Dr. Tonya A. Penkrot
  • Instructor in Anatomy & Physiology, Arizona State University, Phoenix, AZ
  • Ph.D., 2010 - Molar morphometrics and diet in North American condylarths
  • E-mail: tapenkrot(at)

Dr. Kirsten Brown
  • Assistant Professor of Anatomy and Regenerative Biology, George Washington University School of Medicine, Washington, DC
  • Ph.D., 2011 - Obstetrical Adaptation in the Human Bony Pelvis: A Three-Dimensional Morphometric Approach
  • E-mail: brown.kirsten(at)

Dr. Madeleine Chollet
  • Medical Student at Washington University School of Medicine in St. Louis
  • Ph.D., 2011 - The Brain Morphology of Children with Cleft Lip and/or Palate
  • E-mail: cholletm(at)

Dr. Michael Habib

Dr. Frank Varriale
  • Assistant Professor of Biology, Department of Biology, King's College, Wilkes-Barre, PA 18711
  • Ph.D., 2011 - Dental Microwear and the Evolution of Mastication in Ceratopsian Dinosaurs
  • Email: frankvarriale(at)

Dr. Heather Garvin
  • Assistant Professor of Anthropology, Mercyhurst College, Erie, PA
  • Ph.D., 2012 - The Effects of Living Conditions on Human Cranial and Postcranial Sexual Dimorphism.
  • E-mail: hgarvin(at)

Dr. François D. H. Gould
  • Postdoctoral Fellow, Department of Anatomy and Neurobiology, Northeast Ohio Medicalk university, Rootstown, OH
  • Ph.D., 2012 - The Morphology of the Distal Femoral Articular Surface and the Evolution of Cursoriality in Ungulates
  • E-mail: francoisgould(at)

Dr. Evan Garofalo
  • Assistant Professor, University of Maryland, SOM, Department of Anatomy and Neurobiology
  • Research Coordinator, University of Maryland, Shock Trauma & Anesthesiology Research
  • Ph.D., 2012 - Environmental and Genetic Effects on Growth of the Human Skeleton - A Bioarchaeological Investigation
  • E-mail: egarofalo(at)

Dr. Ali Nabavizadeh
  • Postdoctoral Scholar in the Department of Organismal Biology and Anatomy, University of Chicago, Chicago, IL
  • Ph.D., 2014 - Diversity, functional morphology, and evolution of jaw mechanisms in Ornithischian dinosaurs
  • E-mail: alinabav(at)
Dr. Georgina M Voegele
Dr. Katrina Jones

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