Education and Experience
- Harvard Medical School (Boston MA)/ (1957)
- Brigham and Women's Hospital (Boston MA)/ Medicine (1962)
- Mary Imogene Bassett Hospital (Cooperstown NY)/ Medicine/Hematology (1961)
Medicine - Geriatric Medicine and Gerontology
Who''s Who in the East
Who''s Who in Science and Engineering
Who''s Who in America
Who''s Who in the World
American Men and Women of Science
Whos Who in Medicine and Healthcare
UNICEF Gold Medal for East Asia and Pakistan 1983
King Faisal International Prize in Medicine 1984
UNICEF Maurice Pate Prize (ICDDR,B) 1984
W.B. Greenough Award for Excellence in Research/BRAC Student Award 2006
Employee of the Month Recipient/Johns Hopkins Bayview Care Center May 2006
BAFI Outstanding Services Award for the cause of Bangladesh July 2007
- Geriatric Medicine
U.S. Public Health Service 1962-1967 National Heart Institute
Highest rank attained, Senior Surgeon of the National Institutes of Health
Research Associate 1962-1964 Cholera Research Laboratory
Chief Clinical Research 1964-1965 Cholera Research Laboratory
Staff Associate 1965-1967 National Heart Institute
Laboratory of Metabolism
Assistant Chief of Medicine 1967-1970 Baltimore City Hospitals
Assistant Professor of Medicine 1967-1969 Johns Hopkins University
Associate Professor 1969-1983 Johns Hopkins University
of Medicine & Microbiology
Professor of Medicine 1983 Johns Hopkins University
Professor of Microbiology 1983 Johns Hopkins University
Professor of International Health 1985 Johns Hopkins University
Chief, Infectious 1970-1976 Department of Medicine
Diseases Division Johns Hopkins University
Director, Robert Wood 1974-1977 Johns Hopkins University
Clinical Scholars Program
Scientific Director 1977-1979 Cholera Research Laboratory
Director 1979-1985 International Centre for Diarrhoeal
Disease Research, Bangladesh
Faculty member of Division 1985-present Johns Hopkins University at
of Geriatric Medicine and Gerontology Johns Hopkins Bayview Medical Center
1. Shillington JK, Denning GS, Greenough WB III, Hill T Jr, and Ramsay OB. A new method for the resolution of racemic carbonyl compounds; the synthesis and use of 4-(4-carboxy-phenyl) semicarbazide. J Amer Chem Soc. 1958;80:6551-6553.
2. Thomas ED, Lochte HB Jr, Wales M, and Greenough WB III. In vitro synthesis of fetal and adult hemoglobin by fetal hematopoietic tissues. Nature. 1960;185:396-397.
3. Thomas ED, Herman EC Jr, Greenough WB III, Hager EB, Cannon JH, Sahler OD and Ferrebee JW. Irradiation and marrow infusion in leukemia. Arch Intern Med. 1961;107:829-845.
4. Greenough WB III, Thomas ED and Peters T. An intracellular protein intermediate for hemoglobin synthesis. J Clin Invest. 1962;41:1116-1124.
5. Greenough WB III, Sonnenblick E, Januszewicz V and Laragh JH. Correction of hyperaldosteronism and massive fluid retention of unknown cause by sympathomimetic agents. Amer J Med. 1962;33:603-614.
6. Greenough WB III, Gordon RS Jr, Rosenberg IH, Davies BI and Benenson AS. Tetracycline in the treatment of cholera. Lancet. 1964;i:355-357.
7. Benenson AS, Islam MR and Greenough WB III. Rapid identification of Vibrio cholerae by darkfield microscopy. Bull Wld Hlth Org. 1964;30:827-831.
8. McIntyre OR, Feeley JC, Greenough WB III, Benenson AS, Hasan SI and Saad A. Diarrhea caused by non-cholera vibrios. Amer J Trop Med and Hyg. 1965;14:412-413.
9. Lindenbaum J, Greenough WB III, Benenson AS, Oseasohn RO, Risvi S and Saad A. Non-vibrio cholera. Lancet.1965;i:1081-1083.
10. Gordon RS Jr, Greenough WB III and Lindenbaum J. The management of epidemic cholera. Military Med. 1965;130:475-479.
11. Cole JR and Greenough WB III. Patent bile ducts at the time of colourless "rice-water" stool in cholera. Lancet. 1965;ii:972-973.
12. Greenough WB III. Pancreatic and Hepatic hypersecretion in cholera. Lancet. 1965;ii:991-994.
13. Ally KM and Greenough WB III. Electrocardiographic changes during shock due to cholera and other causes. East Pakistan Med J. 1966;10:1-8.
14. Lindenbaum J, Akbar RL, Gordon RS Jr, Greenough WB III, Hirschhorn N and Islam MR. Cholera in children. Lancet. 1966;i:1066-1068.
15. Hirschhorn N, Greenough WB III, Lindenbaum J and Alam SN. Hypoglycemia in children with acute diarrhea. Lancet. 1966;ii:128-133.
16. Rosenberg IH, Greenough WB III, Lindenbaum J and Gordon RS Jr. Nutritional studies in cholera: the influence of nutritional status on susceptibility to infection. Amer J Clin Nutr. 1966;19:384-389.
17. Lindenbaum J, Greenough WB III and Islam MR. Antibiotic therapy of Cholera. Bull Wld Hlth Org. 1967;36:871-883.
18. Lindenbaum J, Greenough WB III and Islam MR. Antibiotic therapy of cholera in children. Bull Wld Hlth Org. 1967;37: 529-538.
19. Greenough WB III, Crespin SR, and Steinberg D. Hypoglycemia and hyperinsulinemia in response to raised free fatty acid levels. Lancet. 1967;ii:1334-1336.
20. Harvey RM, Enson Y, Lewis ML, Greenough WB III, Ally KM and Panno RA. Hemodynamic effects of dehydration and metabolic acidosis in Asiatic cholera. Trans Assoc Amer Physiol. 1968;79:177-186.
21. Harvey RM, Enson Y, Lewis ML, Greenough WB III, Ally KM and Panno RA. Hemodynamic studies in cholera: effects of hypovolemia and acidosis. Circulation. 1968;37:709-728.
22. Enson Y, Harvey RM, Lewis ML, Greenough WB III, Ally KM and Panno RA. Hemodynamic effects of metabolic acidosis in cholera: implications for fluid repletion in severe burns. Ann NY Acad Sci.1968;50:577-583.
23. Judson G, Jones A, Kellogg R, Buckner D, Eisel R, Perry S and Greenough WB III. Closed continuous-flow centrifuge. Nature 1968;217:816-818.
24. Carpenter CCJ and Greenough WB III. Response of canine duodenum to intraluminal challenge with cholera exotoxin. J Clin Invest. 1968;47:2600-2607.
25. Greenough WB III and Buckner D. Removal of microfilariae from unanaesthetized dogs by continuous-flow centrifugation. Trans Roy Soc. Trop Med and Hyg. 1969;63:259-262.
26. Carpenter CCJ, Greenough WB III and Sack RB. The relationship of superior mesenteric artery blood flow to gut electrolyte loss in experimental cholera. J Infect Dis. 1969;119:182-193.
27. Zieve PD and Greenough WB III. Adenyl cyclase in human platelets: activity and responsiveness. Biochem Biophys Res Comm. 1969;35:462-465.
28. Greenough WB III, Crespin SR and Steinberg D. Infusion of long-chain fatty acid anions by continuous-flow centrifugation. J Clin Invest. 1969;48:1923-1933.
29. Crespin SR, Greenough WB III and Steinberg D. Stimulation of insulin secretion by infusion of free fatty acids. J Clin Invest. 1969;48:1934-1943.
30. Carpenter CCJ, Curlin GT and Greenough WB III. Response of canine Thiry-Vella jejunal loops to cholera exotoxin and its modification by ethacrynic acid. J Infect Dis. 1969;120:332-338.
31. Al-Awqati Q, Garcia-Bunuel R, Field M and Greenough WB III. The influence of structural abnormality on ion transport in rabbit ileum. Proc Soc. Exptl Biol Med. 1970;135:598-604.
32. Greenough WB III, Pierce NF and Vaughan M. Titration of cholera enterotoxin and antitoxin in isolated fat cells. J Infect Dis. 1970;121:s111-113.
33. Vaughan M, Pierce NF and Greenough WB III. Stimulation of glycerol production in fat cells by cholera toxin. Nature. 1970;226:658-659.
34. Pierce NF, Carpenter CCJ, Elliott HL and Greenough WB III. Effects of prostaglandins, theophylline, and cholera exotoxin upon transmucosal water and electrolyte movement in the canine jejunum. Gastroenterology. 1971;60:22-32.
35. Hirschhorn N and Greenough WB III. Cholera. Scientific American. 1971;225:15-21.
36. Van heyningen WE, Carpenter CCJ, Pierce NF and Greenough WB III. Deactivation of cholera toxin by ganglioside. J Infect Dis. 1971;124:415-418.
37. Zieve PD, Pierce NF and Greenough WB III. Stimulation of glycogenolysis by purified cholera enterotoxin in disrupted cells. Johns Hopkins Med J. 1971;129:299-303.
38. Al-Awqati Q, Wallace CK and Greenough WB III. Stimulation of intestinal secretion in vitro by culture filtrates of E. coli. J Infect Dis. 1972;125:300-303.
39. Field M, Fromm E, Al-Awqati Q and Greenough WB III. Effect of cholera enterotoxin on ion transport across ileal mucosa. J Clin Invest. 1972;51:796-804.
40. Al-Awqati Q and Greenough WB III. Prostaglandins inhibit intestinal sodium transport. Nature (New Biology). 1972;238:26-27.
Phi Beta Kappa
Alpha Omega Alpha
American Federation for Clinical Research
Infectious Diseases Society of America
American Society for Microbiology
American Society for Clinical Investigation
Fellow of American College of Physicians
Bangladesh Association for Advancement of Science
Bangladesh Medical Society
Association of American Physicians
Fellow, American Association for the Advancement of Science
American Geriatrics Society
Maryland Gerontological Association
Young Investigators Day Committee